In 1989, scientist Hank Pym resigns from S.H.I.E.L.D. after discovering that they attempted to replicate his Ant-Man shrinking technology. Pym believes the technology is dangerous and vows to suppress it.
26 years later, in the present day, Pym’s estranged daughter, Hope van Dyne, and protégé, Darren Cross, have forced him out of his own company. Cross is close to perfecting a shrinking suit of his own, the Yellowjacket, which he shows off to a horrified Pym.
Several months after the events of Avengers: Age of Ultron, well-meaning thief Scott Lang is released from prison after serving a three year sentence. He rejoins his old cellmate, Luis, who tries to recruit Lang into his gang for a burglary, but Lang wants to live an honest life. Lang’s ex-wife is engaged to a policeman named Paxton and will not let Lang see his daughter unless Lang provides child support.
Unable to hold down a job due to his criminal record, Lang agrees to Luis’ offer. Lang breaks into a house and cracks the safe in the basement, only to find nothing but an old “motorcycle suit” that he takes home. Trying it on and pressing a button on it, he inadvertently shrinks to the size of an insect. Terrified by the experience, he returns the suit, but is arrested on the way out. Pym, the homeowner, visits Lang in jail, posing as a lawyer, and smuggles the suit into Lang’s cell to help him break out. Pym manipulated Lang through Luis into stealing the suit as a test, and wants Lang to become the new Ant-Man and steal the Yellowjacket from Cross. Van Dyne has been spying on Cross for Pym, though she still resents her father for not talking about her mother or allowing her to use the suit herself to stop Cross. Pym and van Dyne train Lang to fight and to control ants. They have him steal a device from the Avengers’ headquarters, where Lang gets into a fight with Falcon.
Pym tells van Dyne the truth about the death of her mother: she was Pym’s counterpart, the Wasp, and sacrificed herself to disable a Russian nuclear missile, in the process disappearing into a subatomic quantum realm. Pym warns Lang he could suffer a similar fate if he overrides his suit’s safeguards.
Cross perfects the Yellowjacket suit and invites Pym to the unveiling. Lang, along with his gang and a swarm of flying ants, infiltrates the Pym Technologies building during Cross’ event, sabotages the servers, and plants explosives. When he attempts to steal the Yellowjacket suit, he is trapped by Cross, who had anticipated the theft. Cross, having both the Yellowjacket and Ant-Man suits, sells the technology to Hydra. Lang breaks free of the trap and takes out the Hydra agents. As Cross escapes, the explosives detonates on the chemicals that vaporizes Pym Technologies.
Cross dons the Yellowjacket suit and fights Lang. Lang traps Cross in a bug zapper, but before he can destroy the suit he is arrested by Paxton. Cross takes Lang’s daughter hostage to bait Lang. Lang again fights Cross, and shrinks to subatomic size to penetrate Cross’ suit and sabotage it. Cross is killed by the implosion while Lang uncontrollably shrinks away into the nothingness of the subatomic realm, but Lang perseveres and finds a way to reverse the suit’s mechanism, restoring himself to normal size.
In gratitude for Lang’s heroism, Paxton covers for Lang to keep him out of prison. Seeing that Lang survived and returned from the quantum realm, Pym wonders if his wife is alive, too. Luis tells Scott that the Avengers are looking for him to join them through an intended tip from Falcon.
In a mid-credits scene, Pym shows van Dyne a prototype of a new Wasp costume, and offers it to her. In a post-credits scene, Wilson and Steve Rogers have Bucky Barnes in their custody. Unable to contact Tony Stark due to “the accords”, Wilson states that he knows someone else to contact.
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